Chaetosomodes halli
Acanthocnemidae
A monotypic family from Australia. The type species, the little ash beetle (Acanthocnemus nigricans (Hope)), has been introduced to Southern Europe, Africa, India, Thailand, Myanmar, New Caledonia, and Russia. Known to be pyrophilous in Western Australia, possibly saprophagous. This family is not known from the nearctic region.
(Photo from Kovalenko 2011)
References
Lawrence, J.F.; Leschen, R.A.B. 2010: 9.7. Acanthocnemidae Crowson, 1964. Pp. 262-265 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Chaetosomatidae
14 species in 4 genera: Chaetosoma, Chaetosomodes, and Katachaetosoma from New Zealand; Malgassochaetus from Madagascar. The biology of this family is poorly known, but they are thought to be predators of woodboring beetles and moths. Current phylogenetic work places Chaetosomatidae in an unresolved trichotomy with Metaxinidae and Thanerocleridae. This family is not known from the nearctic region.
(All photos by S.E Thorpe)
References
Leschen, R.A.B. 2010: 9.3. Chaetosomatidae Crowson, 1952. Pp. 247-250 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Chaetosoma scaritides
Mauroniscus maculatus
Mauroniscidae
26 species in 5 genera: Amecomycter (Argenina, Chile, Peru); Mauroniscus (Argentina, Bolivia, Colombia, Ecuador, Peru); Mecomycter (W. North America); Mectemycor (S. California); Scuromanius (S. California, Az/TX south into Mexico, Colombia). The biology of this family is wholly unknown. Not known in Eastern US.
The author advises great caution in spelling with mauroniscid genera names.
(Photos from Coleoptera Neotropical by Juan Enrique Barriga-Tuñón)
References
Lawrence, J.F. 2010: 9.10. Mauroniscidae Majer, 1994. Pp. 271-273 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Majer, K. 1995: Revision of the family Mauroniscidae (Insecta: Coleoptera: Cleroidea). Entomologische Abhandlungen (Dresden), 57: 57-89. (Full revision; Contains keys to all genera)
Metaxinia ornata
Metaxinidae
A monotypic family from the south island of New Zealnd. The type species, Metaxinia ornata, is rarely collected, but is found on living branches and trunks of Nothofagus trees. Insectivourous. Current phylogenetic work places Chaetosomatidae in an unresolved trichotomy with Metaxinidae and Thanerocleridae. This family is not known from the nearctic region.
(Photos by S.E. Thorpe)
References
Kolibáč, J. 2004: Metaxinidae fam. nov., a new family of Cleroidea (Coleoptera). Entomologica basiliensia, 26: 239-268.
Johnson, J.B.; Emberson, R.M.; Marris, J.M.W. 2008: Biology of Metaxina ornata Broun (Coleoptera: Metaxinidae), with notes on associated beetle taxa. Coleopterist's bulletin, 62: 215-219.
Kolibáč, J.; Leschen, R.A.B. 2010: 9.4. Metaxinidae Kolibáč, 2004. Pp. 250-253 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Phloiophilidae
A monotypic family from europe. The type species, Phloiophilus edwardsi, has been reported from Austria, Belgium, Demnark, France, Germany, Hungary, Ireland, the Netherlands, Poland, and the United Kingdom. Larvae are brumal and feed on the fruiting bodies of Phlebia merismoides in Scottland and Peniophora quercina in germany, both of which occur on oak Pupation is vernal to estival. This family is not known from the nearctic region.
(Photo credited to Lech Borowiec and Coleoptera Poloniae)
References
Beetle fauna of Germany family page
Coleoptera Poloniae species page
Crowson, R. A. (1964, December). The habits and life cycle of Phloiophilus edwardsi Steph.(Coleoptera: Phloiophilidae). In Proceedings of the Royal Entomological Society of London. Series A, General Entomology (Vol. 39, No. 10‐12, pp. 151-152). Blackwell Publishing Ltd.
Kolibáč, J. 2008: Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey, 30: 105-133.
Lawrence, J.F.; Leschen, R.A.B. 2010: 9.1. Phloiophilidae Kiesenwetter, 1863. Pp. 239-241 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Phycosecidae
4 decribed species in 1 genus (Phycosecis): P. ammophila (Western Australia), P. hilli (N. Queensland), P. limbata (New Zealand), P. litoralis (Western Australia to New South Wales along southern coastline). Additional, undescribed speciespresent in Vanuatu and New Caledonia. Phycosecids are found on psammolittoral beaches of Australia and New Zealand are coprophagous/necrophagous scavengers. This family is not known from the nearctic region.
(Drawing by D.W. Helmore from Fauna of New Zealand number 37)
References
Leschen, R.A.B.; Beutel, R.G. 2010: 9.8. Phycosecidae Crowson, 1952. Pp. 265-268 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In:Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Prionoceridae
Three extant genera:
Genus Idgia Laporte, 1836 - 130 species throughout Oriental and Afrotropical regions, extending into Asia Minor, India, China, and Japan. Not in Madagascar.
Genus Lobonyx Jacquelin du Val, 1859 - 16 species from Europe, Northern Africa, and Central Asia west to China.
Genus Prionocerus Perty, 1831 - 3 species from India, Taiwan, China, Southeast Asia, and Africa.
Some adults are known to be pollen feeders, while others are saprophagous or predacious.
This family is not known from the nearctic region.
(Drawing of Lobonyx turkestanicus (Kraatz,1882) (Prionoceridae) Modified from Г.Г. Якобсон (1905-1915) "Жуки России": таблица 42.)
References
Leschen, R.A.B.; Lawrence, J.F. 2010: 9.9. Prionoceridae Lacaordaire, 1857. Pp. 268-271 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In:Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Trogossitidae
Three extant subfamilies - Lophocaterinae, Peltinae, Trogossitinae, the later two including approximately 60 nearctic species.
Species level revision, including keys to genera of the world, available in Kolibac 2013.
(Photos by S. E. Thorpe; Grynocharis oblonga, from C. G. Calwer's Käferbuch, Table 14, Picture 20; other drawings from Г.Г. Якобсон (1905-1915) "Жуки России": таблица 19)
References
Kolibac J (2013) Trogossitidae: A review of the beetle family, with a catalogue and keys. ZooKeys 366: 1-194. doi: 10.3897/zookeys.366.6172
Michael A. Ivie (2002). Ross H. Arnett & Michael Charles Thomas, ed. American Beetles: Polyphaga: Scarabaeoidea through Curculionoidea. Volume 2 of American Beetles. CRC Press. ISBN 978-0-8493-0954-0. (incomplete reference!)
Kolibáč, J.; Leschen, R.A.B. 2010: 9.2. Trogossitidae Fabricius, 1801 [sic!]. Pp. 241-247 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Melyridae
Synonymous with Attalomimidae, Gietellidae, Dasytidae, and Malachiidae. The later two are still treated as separate subfamilies by some workers.
Contains the subfamilies Dasytinae Laporte de Castelnau, 1840; Malachiinae Fleming, 1821; Melyrinae Leach, 1815; and Rhadalinae LeConte, 1862.
Byturidae
Until recently, were treated as part of the superfamily Cucujoidea.
Contains the subfamilies Byturinae Jacquelin du Val, 1858 and Platydascillinae Pic, 1914
Biphyllidae
Until recently, were treated as part of the superfamily Cucujoidea.
References:
Goodrich, M.A. and Springer, C.A. 1992. A revision of the family Biphyllidae (Coleoptera) for America, north of Mexico. The Coleopterists' Bulletin 46:361-377.